Scotland coach Matt Williams has made seven changes, including two positional switches, for the opening RBS 6 Nations Championship match against France in Paris on Saturday.
Hugo Southwell has moved from full-back to inside centre with Chris Paterson taking on the number 15 role.
Other changes made following the 45-10 loss to South Africa in the last Abbey Autumn Test see Ben Hinshelwood, Andrew Henderson and Nathan Hines lose starting places, while Donnie Macfadyen and Allan Jacobsen are discounted through injury.
Centre Andy Craig returns for the first time since Scotland's 2003 World Cup pool defeat by France; Simon Danielli, Tom Smith and Scott Murray are back after injury; and Jon Petrie will start at openside in the back-row alongside Allister Hogg.
Scotland team: C Paterson (Edinburgh); S Danielli (Borders), A Craig (Glasgow), H Southwell (Edinburgh), S Lamont (Glasgow); D Parks (Glasgow), C Cusiter (Borders); T Smith (Northampton), G Bulloch (Glasgow, captain), G Kerr (Leeds), S Grimes (Newcastle), S Murray (Edinburgh), J White (Sale), A Hogg (Edinburgh), J Petrie (Glasgow).
Replacements: R Russell (London Irish), B Douglas (Borders), N Hines (Edinburgh), J Dunbar (Leeds), M Blair (Edinburgh), G Ross (Leeds), B Hinshelwood (Worcester)
Before Will Greenwood started breaking down moves off the field, he was doing the business on it - and no match better illustrated the type of marauding centre he was than in a virtuoso performance against Wales.