The British and Irish Lions won their first Test match for eight years after a stunning end-of-tour 22-9 performance at Ellis Park left world champions South Africa reeling.
Shane Williams' try double and a second-half interception score by his fellow wing Ugo Monye - plus 13 points from fly-half Stephen Jones - sent South Africa crashing to a rare defeat on the ground in Johannesburg where they won the 1995 World Cup.
The Lions ended a run of seven successive Test losses - they had not triumphed since beating 2001 hosts Australia in Brisbane - and also avoided a first series whitewash at South Africa's hands.
And the result saw them match the record 28-9 Lions success of Willie John McBride's all-conquering 1974 tourists in Pretoria.
The Lions had heroes everywhere, no-one more so than number eight Jamie Heaslip, who provided a towering presence in all areas, while centre Riki Flutey, flanker Martyn Williams and scrum-half Mike Phillips weren't far behind.
Stephen Jones missed an early 40-metre penalty chance for the Lions, but the opening exchanges proved every bit as brutal as last Saturday's Loftus Versfeld classic, with both teams tearing into each other.
And it took the Lions just nine minutes to strike a clear psychological blow when prop Phil Vickery, dropped after his first Test mauling at the hands of Springboks scrum destroyer Tendai "The Beast" Mtawarira, responded by spearheading a huge Lions scrum.
Springboks hooker Chiliboy Ralepelle popped out of his binding, and Jones kicked the Lions ahead before Morne Steyn slotted an equalising penalty, his first of three in the game.
Phillips made a colossal tackle on Springboks centre Jaque Fourie - it forced him to temporarily make way for substitute Francois Steyn - before flanker Joe Worsley pinned home skipper John Smit with a crunching hit.
And as South Africa regrouped, the Lions struck with a smartly-taken try.
Heaslip galloped into space, and the delivered a neat inside ball to Williams, who claimed his first try of the tour in probably his final Lions appearance.
Jones should have added an easy conversion, but the the ball fell off his kicking tee as he ran up, and his flailing attempt to send a drop kick over failed.
There was no let-up in the ferocity, but the Lions struck another telling blow nine minutes before half-time.
It was a try of breathtaking brilliance, all created by Flutey, whose kick over retreating Springboks wing Odwa Ndungane, ended with him astonishingly flicking the ball inside one-handed and Williams sprinted over unopposed.
Jones converted for a 15-3 advantage as the interval approached, and no-one could deny the Lions a handsome 12-point cushion, highlighted by some devastating finishing.
The Lions though, had lock Simon Shaw sin-binned two minutes before the break for use of the knee on Springboks scrum-half Fourie du Preez.
And Steyn slotted a second penalty on the stroke of half-time, cutting the deficit to nine points.
De Villiers made a double substitution for the second period, sending on Ruan Pienaar and Bismarck du Plessis, yet it made no immediate difference as the Lions powered on in search of that elusive Test triumph.
But centre Tommy Bowe was then forced into frantic defensive mode, preventing a certain try by knocking the ball out of debutant Springboks full-back Zane Kirchner's hands.
South Africa kept pressing, only for centre Wynand Olivier to see a floated pass intercepted by Monye 70 metres out, and the England speedster collected his fifth try on tour.
Jones converted, putting the Lions in dreamland at 22-6 clear.
The Lions lost an injured Flutey - scrum-half Harry Ellis replaced him and Phillips moved into midfield - but they weren't to be denied as two more Jones penalties rubbed South Africa noses in the dirt before Ndungane was denied a consolation try by the television match official.
As head coach Stuart Lancaster polishes the axe he will soon wield over his World Cup training squad, Exeter Chiefs coach Rob Baxter has backed Jack Nowell, Henry Slade and Luke Cowan-Dickie to be given a stay of execution.
Italy Under-20s held off a late Samoa blitz to retain their place in the World Rugby U20 Championship on Saturday and head coach Alessandro Troncon believes the experience will have served as a valuable lesson for his young troops.