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Ashton has scored three tries in three appearances against Australia – including a memorable effort at Twickenham two years ago – but head coach Stuart Lancaster insists it’s the team ethos that’s most important.
He said: "To select 23 from a squad of 30 plus who have trained so well, with such intensity and focus, was difficult but it's a good position to be in. The "team first" mentality of all our squad has impressed us greatly.
"We were pleased with lots of aspects of our performance last week, but we know we need to improve this week and, by having some consistency in selection, we allow those players to demonstrate we can step up to the next level against what will be a very tough Australian side.
"The crowd at Twickenham last week were brilliant and we can't wait to get back there this Saturday and take part in what I am sure will be a very exciting Test match.”
Alex Goode continues at full-back after impressing in the 54-12 victory over Fiji with Brad Barritt and Manu Tuilagi again completing the midfield partnership.
Toby Flood and Danny Care again start at half-back while Joe Marler, Tom Youngs – who won his first cap against Fiji – and Dan Cole complete the front row.
Geoff Parling and Tom Palmer line up in the second row while Tom Johnson plays at blindside, Chris Robshaw continues to lead the side from openside and Tom Waldrom keeps his place at No.8.
ENGLAND:
15. Alex Goode
14. Chris Ashton
13. Manu Tuilagi
12. Brad Barritt
11. Charlie Sharples
10. Toby Flood
9. Danny Care
1. Joe Marler
2. Tom Youngs
3. Dan Cole
4. Tom Palmer
5. Geoff Parling
6. Tom Johnson
7. Chris Robshaw
8. Thomas Waldrom
Replacements:
16. David Paice
17. David Wilson
18. Mako Vunipola
19. Joe Launchbury
20. Tom Wood
21. Ben Youngs
22. Owen Farrell
23. Mike Brown