The Newport Gwent Dragons forward is suffering from vertigo, and could not join his fellow World Cup contenders on the flight to Brittany.

Wales coach Gareth Jenkins said: 'Colin is suffering from viral labyrinthitis, or vertigo, which causes imbalance and nausea, making him too unwell to travel with us on Saturday.

'Initially, we thought Colin would join us for the remainder of the week, but he is still unable to fly at this stage so we felt it was more important that he makes a full recovery in order to resume full training on our return from France.

'Other than Colin, everyone has taken an active part in training.

'Ian Evans, Rhys Thomas and Gavin Henson, who are recovering from injury, have taken a full, active part in training, with Shane Williams resuming all training except contact work.

'Jamie Robinson suffered a minor ankle ligament strain yesterday (Sunday), and he has been resting today.'

Wales are based at their World Cup training centre near La Baule ahead of facing England in a tournament warm-up Test at Twickenham on Saturday week.

Jenkins added: 'It is great to have this week to concentrate on rugby training after a period of conditioning.

'The focus is on contact work and on preparation for the England match.'