The return to fitness of Morrison – impressive during the 2008 RBS 6 Nations – means another Scotland centre, Andy Henderson, drops to the bench.

Lome Fa’atau misses out due to a fractured shinbone so Hefin O’Hare moves to the right wing with Scotland winger Thom Evans on the left.

In the front row, Scotland hooker Dougie Hall is recalled in place of another Test player, Fergus Thomson, despite being concussed against Bath while prop Ed Kalman edges out Moray Low.

Finally, captain Alastair Kellock, in the Scotland squad for the Autumn Tests, returns to replace of Dan Turner, who has to settle for a place on the bench.